the button displays the rhythmic value of the current melody note (assuming that note was added to the chords table). When clicked upon, the program switches to Play mode ready to change that rhythmic value to a different one.
This is used occasionally because the normal way to setting the rhythm of a song is to set the rhythmic value of all melody notes in the Play mode. Not by switching back and forth between the Melody mode and the Play mode.
See The Best Way to Enter Full Songs.
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